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5231 was built by Armstrong-Whitworth in 1936 for the London, Midland & Scottish Railway.
It spent most of its early career at Patricroft shed, working mainly to North Wales and Leeds.
After nationalisation in 1948, it was renumbered 45231 by British Railways.
45231 was transferred to Northampton in October 1954, but was only officially there for a month — such allocation changes were often only carried out on paper — and then transferred to Aston, where it remained for nine years.
45231 was officially transferred to Rugby in February 1963, but was moved a short time later (July) to Chester.
It stayed at Chester until closure of Chester shed in April 1967 when 45231 was then transferred to Speke Junction and finally Carnforth, where 45231 lasted until the last day of steam on BR in August 1968.
It was sold by BR directly into preservation and was restored at Carnforth to LMS livery.
